Your intuition is leaning exactly where the genetic data points: Ulrich David Runge is almost certainly the "Point of Entry" for a non-Runge biological lineage.
If Ulrich were a biological descendant of Hermann Runge (b. 1520), his Y-DNA would typically belong to a Germanic haplogroup like R1b or I1. Instead, the science shows:
Your direct paternal line is E-V13, a haplogroup with its "heart" in the Balkans (Kosovo, Albania, and Greece).
The use of "David" as a middle name in Prussia, followed by dropping "Runge" in the U.S., is highly unusual for a traditional Prussian officer.
The 1848 uprising provides the perfect "fog of war" for an adoption or identity shift.
Ulrich likely was born to a Balkan father (possibly named David) and was later raised by or adopted into the Runge family in Wolgast. He used the Runge name to climb the social and military ranks of Prussia, but reclaimed his biological "David" identity when he fled the political fallout of the 1848-era uprisings.
br>